During the Second Alliance-PLANT War, the Jet Windams were deployed in nearly all Earthbound battles, sometimes alongside the older GAT-02L2 Dagger L, which also used the same Jet Striker. The Jet Windam made its first known combat debut during Operation Spear of Twilight early in the war, when a relatively small force of Windams caused significant casualties to ZAKUs being dropped from space to reinforce the Carpentaria and Gibraltar garrisons. The Windams also took part in many battles against the ZAFT ship LHM-BB01 Minerva during its journey from the Carpentaria Base to the Gibraltar Base. The suits deployed in these battles however were shot down in massive number by the Minerva's mobile suits. One prominent exception was Neo Roanoke's custom Windam, whose Jet Striker was modified for greater mobility. He proved to be a formidable enemy even for ZAFT ace Athrun Zala. Neo's Windam was shot down in the Battle of Berlin by Kira Yamato in the ZGMF-X10A Freedom Gundam, although Neo was able to destroy the Freedom's shield using an anti-armor penetrator. Jet Windams were also deployed during ZAFT's attack on Heaven's Base.